Leptodactylid Frogs of the Physalaemus pustulosus Group

Abstract
Within the leptodactylid frog genus Physalaemus, 4 spp. comprise a monophyletic group: P. petersi (with P. freibergi designated as a junior synonym), P. pustulatus, P. pustulosus and P. coloradorum, sp. nov. from the Pacific lowlands of Ecuador. P. petersi and P. pustulosus seem to be sister-species, as do P. pustulatus and P. coloradorum. The monophyly of the genus Physalaemus is questionable.