Abstract
The epidermis of Anemarrhena asphodeloides is composed of three distinct cell types: guard cells of the stomata, ordinary epidermal cells, and papilla cells occurring in files wedged between files of ordinary epidermal cells. Both stomata and papilla cells have their origin from formative mitoses in the young protoderm. Their differentiation described on the basis of light and polarization microscopy involves a decrease in cell contacts and changing orientation of cellulose microfibrils of the cell wall. These changes are discussed in relation to the pattern of cell divisions and to the modification of cell shape during epidermal development.