Abstract
Evoked and spontaneous long-distance vocalizations of wild coyotes (Canis latrans) were studied in Texas and Oklahoma from June 1972 to August 1974. Sonagrams of tape recordings showed that adult coyotes produced two basic sounds—the bark and the flat howl. These sounds were given by animals in both single and group situations. In addition the yip, yipe, short-howl, warble, laugh, and irregular howl were common group vocalizations. Two or more sounds were often joined to form combination howls. The scream and gargle were characteristic of immature coyotes. Sound patterns from the bark to the flat howl formed a continuum of increasing phonetic complexity with intermediate sounds frequently produced. Coyote vocalization was variable in sound type, duration, Hertz, amplitude of fundamental and other harmonics, and in the sequence of combinations of two or more different sounds. In these ways information could be conveyed.

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