Refugee communities of Carbonicola

Abstract
Sparse elongate forms of the non-marine bivalve Carbonicola are associated with an abundance of trace fossils comprising resting traces (Cubichnia) referable to Lockeia, trails (Repichnia) and escape shafts (Fugichnia), all in arenaceous sediments of the Upper Carboniferous. Associated organic-rich argillaceous sediments contain the well-known, abundant and highly variable Carbonicola body fossil faunas. The latter are interpreted as the remains of shells preserved in what was the typical life environment (characterized by muddy sediments and slow rates of sedimentation). The elongate shells associated with escape shafts are the only form of Carbonicola capable of escape upwards from burial by rapid sedimentation. For such fugitives from preexisting established communities, the term refugee community is proposed.

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