Influence of Adult Females on within-Shelter Distribution Patterns of Blattella germanica (Dictyoptera: Blattellidae)

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Abstract
The spatial distribution of a mixed-age group of German cockroaches within four adjoining shelters is investigated. Experimental variables are female density and female reproductive state. There is an inverse relationship between overall population aggregation and female density if the females lack oöthecae. If oöthecae are present, aggregation increases with density, with a more intense aggregation if the females carry immature rather than mature öthecae. The inter- and intracomponent aggregation relationships underlying these patterns are examined. Cyclic production of aggregation pheromone and possible production of a repellent by adult females are suggested. This has implications in respect to the evolution of interrelated behavioral characteristics.

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