Kondous laventicus, a New Ceboid Primate from the Miocene of the La Venta, Colombia, South America

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Abstract
A new genus and species of fossil primate, Kondous laventicus, part of the Miocene La Venta primate fauna of Colombia, South America, is described here for the first time. Screening at a new quarry within the Monkey Unit of the Honda Formation, the Kyoto Site, has revealed dental remains of this new form of a large-sized ceboid. The La Ventan vertebrate fauna is comparable to the Friasian Land Mammal Age, indicating a middle Miocene age. Biostratigraphic studies suggest a 13.6-15.2 Ma for the Friasian faunal zone. Kondous is most closely allied to extant spider monkeys, Ateles, providing the first fossil evidence of their phylogeny. It also implies a nearly three-fold discrepancy between the molecular and paleontological estimates of the age of the Ateles lineage.