A statistical study ofRhamphorhynchusfrom the Solnhofen Limestone of Germany: Year-classes of a single large species

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Abstract
A statistical and morphological study of the pterosaurRhamphorhynchusfrom the Solnhofen Limestone of Germany shows that specimens fall into discrete size-classes that are year-classes resulting from seasonal mortality or preservation of specimens. Taxonomic characters used in the past to separate species ofRhamphorhynchusare all related to size and ontogeny, and all specimens belong to a single species,R. muensteri.Thus, the collections ofRhamphorhynchusprovide an unequaled record of the growth and development of a Jurassic rhamphorhynchoid pterosaur.Rhamphorhynchusdid not have rapid determinate growth; the growth rate was comparable to that of extant crocodilians, and growth continued over the course of at least three years after individuals began to fly.