The nuclei in hyphae, oospores, and gametangia of Achlya klebsiana and Saprolegnia ferax were examined by light microscopy after stain fixation with acetoorcein. It was found that the vegetative phase is diploid and that meiosis is not zygotic but is gametic in these organisms. The nuclear divisions in hyphae and germinating oospores were similar and were classically mitotic.