Postfertilization Wandlabrinthe Formation and Function in the Central Cell of Soybean, Glycine max (L.) Merr. (Leguminosae)

Abstract
A Wandlabrinthe develops around the micropylar end of the central cell at the time of ovule maturation and fertilization in soybean, G. max (L.) Merr. Wandlabrinthe development occurs during morphogenesis of the central cell as it is transformed from being 1 of the 7 cells of the megagametophyte (prefertilization state) to becoming the embryo sac after syngamy (postfertilization state). The Wandlabrinthe circumvallates the synergids by forming a ridge of cell wall material that projects into the central cell toward the zygote and the synergids at the level of the filiform apparatus. Degenerating nucellar cells are adjacent to, and contiguous with, the Wandlabrinthe, and cells in this region of both integuments are rich in starch reserves. The Wandlabrinthe most likely functions in translocating nutrients from degenerating nucellar cells into the central cell/embryo sac for endosperm and embryo nutrition.