Sur la morphologie des akènes de quelques Borraginacées de la flore de France. Intérêt systématique de cette étude

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Abstract
Among the Boraginaceae the achenes generally number 4, thus forming a typical tetrachene. An exception is the genus Cerinthe where these 4 elements are grouped in such a way that the fruit appears externally divided into 2 parts (diachene). Abnormal or incomplete development is observed as a result of the inadequate formation of 1 or several achenes. In certain Heliotropioideae (i.e., Heliotropium supinum L.) 3 out of 4 achenes frequently abort; among the Eritrichiae there are often fewer than 4 achenes in the genus Eritrichium; among the Anchusa, 1 or 2 achenes in the genus Alkanna. The morphologic characteristics of the achenes of 10 genera are listed in detail. In the Boraginaceae as among the Labiatae, the achenes can be a useful adjunct in determining the identity of the plants. This is particularly true with Omphalodes, Cynoglossum, Echinospermum, Symphytum, Anchusa, Nonnea, Alkanna, Myosotis, Lithospermum and Echium.

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