Asymmetric, two-sided competition in even-aged monocultures of red pine

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Abstract
A framework for analysis of even-aged plant monocultures was developed and used to examine hypotheses on the process of competition, using data obtained from three independently established red pine (Pinus resinosa Ait.) stand-density experiments. Growth rate and relative production rate (a measure of tree vigour) were computed for the individual trees in stands of differing density at points in time when these stands had the same mean stem volume. Competition is shown to be both asymmetric (small trees lose vigour more rapidly than large trees as density increases) and two-sided (not only do larger trees affect smaller trees, but to some degree small trees also cause growth reduction in larger neighbours). Increasing density results in increased size inequality as smaller trees lose vigour and become suppressed. The asymmetric, two-sided competition process in red pine stands is considered ecologically inherent for a species adapted to existence in even-aged monocultures.