Agra, arboreal beetles of Neotropical forests: mixta group, virgata group, and ohausi group systematics (Carabidae)

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Abstract
This paper is the fifth in a series designed to cover taxono‐mically all species‐groups of the lebiine genusAgra, whose cumulative ranges extend from southernmost Texas to northernmost Argentina. Themixta, virgata, feisthameliandohausigroups constitute Section Feisthameli of the genus, and a key to these groups is provided. Themixtagroup consists of two species which have a composite range extending from southeastern Brazil to middle Mexico; thevirgatagroup consists of nine species which have a composite range extending from southeastern Brazil to Guatemala; and theohausigroup consists of four species which have a composite range restricted to southeastern Brazil. Thespeciose feisthameligroup is covered in a separate paper.Among the three groups in the present paper, eleven new species are described from the following type localities:perkinsorum(MEXICO, Nayarit, 17.6 km southwest of Compostela),vate(MEXICO, Veracruz, Los Tuxtlas, La Playa Escondida),nola(REPUBLIC OF PANAMA, Panama, Cerro Campana),cadabra(EQUADOR, Pichincha, 47 km south of Santo Domingo de los Colorados, Rio Palenque Station),chocha(WEST INDIES, Trinidad, Arima Valley),phainops(FRENCH GUIANA, Pariacabo),imaginis(BRAZIL, Goias, Jatahy),inca(PERU, Madre de Dios, Rio Tambopata Reserve, 30 km (air) southwest of Puerto Maldonado),itatiaya(BRAZIL, Rio de Janeiro, Parque Nacional Itatiaya),calamitas(BRAZIL, Bahia, Itamaraju),anthrax(BRAZIL, Espirito Santo, Parque Socretama). A dot map illustrates the range of each taxon.Distribution and relationships are discussed in general, but detailed cladistic and biogeographic analysis is deferred until taxonomy of the four groups of Section Feisthameli is completed. Species distribution correspond to Pleistocene refuges suggested by other authors based on other groups and classes of organisms.