Abstract
A partly complete skeleton from the late Paleocene of western North Dakota is described as Simoedosaurus dakotensis, n. sp., and is compared with European individuals of the genus. This specimen represents the earliest occurrence of the genus and is the first species to be recognized from North America. Skull characters that differ from the type species and indicate an early stage of development are: shorter muzzle; marginal teeth of more uniform size; larger lachrymal and a broader and longer skull table. Postcranial distinctions from its close ally Champsosaurus, with which it is often confused, are also presented, and its association with crocodilians is noted.