On the Origin of Mating Behavior in Spiders

Abstract
In the mating of spiders spermatic fluid is first deposited before being taken up by the palpal organs of the male for introduction into the female. From a comparative survey of the mating habits of other arachnids, it is suggested that ancestral spiders transfered sperm to the female by way of a spermatophore deposited on the substratum, but that, with the development of a web-spinning habit, the spermatophore was lost.

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