Abstract
A new species of sawfly, Cladomacra terricola, belonging to the family Pergidae, is described from an apterous female from mid‐montane Papua New Guinea. The profoundly modified thorax of the female is compared with the thorax of other wing‐reduced and macropterous Hymenoptera. The ovipositor is greatly reduced and lacks the typical, symphytan, saw‐like valves. The ordinal, family, subfamily and generic classification of C. terricola is discussed. Diagnostic features of the pergid subfamilies Pterygophorinae, Euryinae and Perreyiinae are reviewed. The egg and larva of C. terricola are described. Egg brooding by the female is reported. Larvae of C. terricola may feed on dead plant material in forest litter.