On the Attachment of Non-Homologous Chromosomes at the Reduction Division in Certain 25-Chromosome Daturas

Abstract
The primary 2n + l mutants, prising from the addition of 1 chromosome to the 2n group, may form tertiary 2n + l mutants, after crossing with an iso-morphic strain of 2n plants which gives abnormal Mendelian ratios with regard to certain genes in the crosses. The tertiary mutants are distinguished by certain non-homologous chromosomes being combined into trivalents or quinquevalents at the maturation divisions. From the nature of the chromosome configurations, and from the results of measurements, it is presumed that the isomorphic strains contain 2 pairs of chromosomes resulting from segmental interchange between non-homologous chromosomes.

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