Leclercqia, Haskinsia, and the Role of Leaves in Delineation of Devonian Lycopod Genera

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Abstract
Leclerqia, Protolepidodendron, and Haskinsia are shown to be nearly identical when leaves are absent or incompletely preserved. Hence, leaves are essential to positive identification of the three genera. The sporangia of Leclerqia are sessile and may have a tapetal layer and well over 1,000 spores each. Stomata on leaves, sporophylls, and sporangia are sunken. Additional dichotomies of the five-tipped leaves occur. All dichotomies are at right angles. Leclerqia is eusporangiate, homosporous, and Haskinsia grew is suggested. Reconstructions and emendations of both genera are presented. Phylogenetic relationships among Devonian lycopods are discussed briefly.