Plasmatic Filaments and Particles in Mature Sieve Elements of Heracleum sphondylium under the Electron Microscope

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Abstract
By using dimethyl sulphoxide in addition to glutaraldehyde on the phloem of Heracleum a speedier fixative for phloem sieve elements has been developed, less disruptive to plasmatic filaments (p. f.)- Strands of p. f. under the electron microscope seem to run from sieve element to element, axially through the lumina and the sieve-plate pores. Some of these axial strands give profiles which suggest that they exist as tubular structures in which a series of swellings, 20 to 60 ran in diameter, can be detected as if microperistalsis were passing along them. Marker particles of five different types were noted, often attached to wisps of plasmatic filaments. Many of the p. f. seemed to have a helical substructure.

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