Consideration is given to chromosome behavior at the reduction division in Uvularia, with especial emphasis upon a comparison between conditions in plants and animals. Material was prepared by the iron-acetocarmine method and examined under a water-immersion objective. Differences in size and form make it possible to identify the chromosomes individually. The tetrads are relatively simple in structure [long dash]less complicated than those found in some insects. Evidence is summarized as follows (1) There is apparently fusion at the junctions of the 2 constituents of each bivalent. (2) Horizontal rings and V''s divide into their constituent ehromatids along a horizontal plane, the line of division being distinct at the metaphase. (3) Vertical V''s and rings have not been observed in the act of separating } but they probably separate into upper and lower halves. (4) The rings and V''s apparently diminish as the loop between them is pulled out by the spindle fibers. This would presuppose crossing or interlacing of the chromatids.