Center-Edge Effect in Red Squirrels: Evidence from Playback Experiments

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Abstract
Larder-hoarding red squirrels (Tamiasciurus hudsonicus), with a valuable resource (midden) within their territory, should respond more intensely to intrusion at the center than on the edges of their territory. In a playback experiment, squirrels gave more aggressive screech calls in response to simulated intrusion at their midden than at the territory edge, but moved more often in response to simulated intrusion at the edge. The frequency of territorial rattle calls and of alarm calls did not change as long as playback was from within the territory.

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