Abstract
In F1 plants from crosses between T. aestivum cv. ''Chinese Spring'' monosomic 5D and A. speltoides Tausch. var. aucheri Boiss at a temperature of 12.degree. C the presence of chromosome 5D is necessary to maintain the level of homoeologous pairing and chiasma formation observed under greenhouse conditions. Some genotypes of A. speltoides, however, were more efficient than others in compensating for the absence of chromosome 5D. The differences in compensating ability indicate the existence of several alleles in A. speltoides that counteract the low temperature pairing system on chromosome 5D. An analogous system of alleles previously reported in A. speltoides that suppresses the chromosome 5B diploidizing system of wheat was substantiated. The 2 systems are independent.