Trophic complexity in a littoral boulderfield

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Abstract
A food web constructed for the major invertebrate carnivores in a littoral boulderfield suggests that trophic complexity was very low. Each of the predators consumed several prey species but monopolized one or two species, producing a sharp partitioning of food resources. Spatial heterogeneity and differences in feeding behavior reinforced this pattern. The distributions of bored shells of Protothaca staminea and Tegula funebralis among the size classes of empty shells available in the boulderfield suggested that their primary predator, Octopus, had no size preferences.

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