Predictions as to Chromosome Configuration, as Evidence for Segmental Interchange in Oenothera

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Abstract
The correctness of the segmental interchange theory, as applied to Oenothera, is tested by a method which involves making predictions, based upon this theory, as to the chromosome configurations to be found in certain hybrids. The arguments upon which a total of 37 predictions were made are recounted. All of these predictions were later found to be correct, proving the correctness of the segmental interchange theory. Establishment of this theory in turn proves that paternal and maternal chromosomes alternate in the circles formed in diakinesis, for each chromosome must be adjacent to those chromosomes which have ends homologous with its own.