Abstract
Native perennial herbs [Solenomelus peduncularis and Pasithea coerulea] in central Chile tend to be most abundant beneath bushes. This may be because the humid microclimate under bushes prevents desiccation (the absence of herbs from open spaces would then be explained by water stress), or it may be because herbivores which eat the herbs forage more in the areas between rather than under the bushes. The problem was studied experimentally by removing half the canopy of 8 individual bushes of the same species, and then preventing grazing of the herbs in 1/2 of the removed section with an exclosure. This design allowed comparison of areas protected against desiccation and herbivores (the uncut half of the shrub), with areas protected against herbivores but not desiccation (inside the exclosure) and with unprotected areas exposed to both desiccation and grazing. Grazing (most likely by introduced rabbits) was probably the main cause of the scarcity of native herbs between bushes and of their relative abundance beneath bushes.

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