Abstract
The dominant gene C controls the production of anthocyanin pigmentation on the inner side of the corolla limb. On a white background, the color is frequently Pomegranate Purple, Amaranth Purple or Rose Color, but if the background is orange the color is frequently Carmine or Ox-blood Red. The exact color depends also upon unidentified modifying genes; it is often impossible to determine from the color of the anthocyanin whether the background is white or orange. The genes splotched, granular, red-outline and bicolor are all recessive and are expressed only in the presence of C. Splotched (sp) restricts the anthocyanin pigmentation to irregular blotches and streaks, is apparently epistatic to granular, and is linked with C with about 20% crossing over. Granular (gr) produces numerous small colorless dots in the anthocyanin pigmentation. Red-outline (ro) restricts the anthocyanin on the lower lips to an outline with occasional small patches of color. Bicolor (bi) produces a flower with Pomegranate Purple upper lip and Rose Color lower lip, and is linked with the self-sterility alleles with about 34.6% crossing over.

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