The Estimation of Sampling Biases for Male Tsetse

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Abstract
A method is suggested for estimating sampling biases for male tsetse, based on the assumptions of the classical feeding cycle model. Sampling intensity for any particular method is defined as the catch of flies in different segments of the non-feeding phase of their current cycles, expressed as a percentage of the catch of flies that are just entering the feeding phase. Two examples are given, of G. palpalis in Nigeria and G. morsitans in Zambia. In both cases, sampling intensity is low just after feeding and increases slowly at first, and then rapidly as flies enter the feeding phase. Flies in their feeding phase can apparently find and feed on hosts within 1 day. Feeding rates are inversely related to the fat content of flies entering the feeding phase of their current cycle; rates of fat usage increase with this variable. It is important to distinguish between tsetse flight activity in general, and feeding behavior in particular. It is emphasized that the analytical technique proposed depends on the still inadequate understanding of the field behavior of flies.