Abstract
The fore wing bases of Monotrysia can be homologized with the plan for a generalized insect wing base (Snodgrass, 1935). The wing bases of advanced Lepidoptera are very different from those of primitive forms. An evolutionary series can be traced through the order showing how the changes may have taken place.The positions of the bending cuticle areas vary very little within the order. Within each superfamily the axillary region is surprisingly constant. It is impossible to distinguish between the wing bases of many related ditrysian families.The jugum is particularly interesting; it is present in all the members of the old group “Microlepidoptera” and not only in the Monotrysia or “Jugatae”.

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