Genetic Meaning of Zooecial Chamber Shapes in Fossil Bryozoans: Fourier Analysis

Abstract
Fourier harmonic amplitudes quantitatively characterize chamber shapes of fossil tubular bryozoans. The odd-numbered harmonics, particularly the seventh, carry evolutionary information. The phenotypically plastic second and sixth harmonics measure zooecial orientation and packing, respectively. As a measure of crowding, the sixth harmonic reflects mechanistic growth response to paleoenvironmental conditions.