Acanthodian and coelacanth fish from the Dinantian of Foulden, Berwickshire, Scotland

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Abstract
Acanthodians are represented by numerous young and poorly cranially ossifiedAcanthodes ovensiWhite, the earliest known acanthodid. Study of the smallA. ovensitogether with the type series shows that squamation begins posteriorly, and spreads anteriorly along an initially triangular front, as inA. bridgeiZidek.Gyracanthussp. occurs at several horizons in the sequence. The coelacanth remains include the first articulated specimen known from Foulden, but preservation only allows an assignment to cf.Rhabdoderma. Although these taxa are distinctive, resemblances to other taxa occur only in generalised features. Hence, unlike the Foulden actinopterygian fish, the acanthodians and coelacanths cannot be used to suggest closer affinities with any other fish of this age, on which information is sparse.

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