Abstract
Sporelings of Marsilea, grown aseptically in various concentrations of sugars, particularly glucose, have been investigated anatomically. It is shown that, corresponding to the features of the external morphology, the anatomy of the sporelings from the lower sugar concentrations (1 and 2 per cent. glucose) has many characteristics of the water forms of amphibious plants, while that of the sporelings from the higher sugar concentrations (4 and 5 per cent, glucose) has the characteristics of typical land forms. A number of the differences between the two types of sporeling may be referred to a change in the duration and direction of cell-division in the rhizome-apex. Views as to the factors influencing the origin of land or water forms are briefly discussed.

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