Abstract
Hyphae of H. fuscum invade alder wood and attack all cell types. Eventually, hyphae emerge into the bark, rupture the periderm, and from a stroma. Coiled ascogonia are produced in the stroma. Each ascogonium becomes ensheathed in a hyphal envelope which ultimately develop into the ascocarp wall. The ascogonium becomes segmented, falling apart into cells. Some ascogonial cells produce ascogenous cells; others become inflated and branched. Polar growth of the envelope and filamentous space-making hyphae are involved in peri-thecial expansion. Developing asci grow into the mass of space-making hyphae and lyse some of it. The developmental pattern is of the Xylaria type.