Revision of the Genus Joosia (Rubiaceae)

Abstract
Seven species ofJoosia are recognized, four of which are new:J. ulei,J. aequatoria,J. standleyana, andJ. macrocalyx. All of the species are confined to western South America from Colombia south to Peru with the main center of dispersal in Ecuador. A new subgenus,Sectocalyx, is also described.

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