Seasonal Shifts in the Spawning Site of a Northeast Pacific Intertidal Fish

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Abstract
In Barkley Sound, Vancouver Island, Canada, the spawning of the black prickleback, Xiphister atropurpureus, occurs during winter on rocky beaches protected from wave action. However, during spring, spawning occurs on rocky beaches of increasing exposure. This species spawns only under boulders with pebble, small rock, or shell substrates. Coincident reduction of wave action and increases in temperature probably cause the seasonal shift in the spawning site of this stichaeid.

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