Chemical Changes inLolium temulentumL. after Treatment with (2-Chloroethyl) Trimethylammonium Chloride (CCC)

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Abstract
Two experiments with an inbred line of the summer annual Lolium temulentum L. are described. The plants were grown under 18-hour days in 8.5 cm pots containing pure vermiculite. The first experiment was concerned with the effect of (z-chloroethyl) trimethylammonium chloride (CCC) upon soluble carbohydrate content, and the second with the combined effects of CCC and nitrogen upon soluble carbohydrate, free α amino-N, and chlorophyll content. Concentrations of CCC from 0.01–0.5 M inhibited growth and induced large increases in the free sugar content of the plant. With 0.05 M CCC and above, large amounts of fructosan were formed. Adequate nitrogen supply lowered the free sugar level and large amounts of free α amino-N appeared. Crude protein content (per cent dry weight) was increased by CCC at low nitrogen levels but was less affected when N was adequate. Chlorophyll production was stimulated in the presence of CCC. The metabolic implications of the CCC-induced chemical changes are discussed and a possible scheme for CCC action is suggested.