Long-term compositional dynamics of Acadian mixedwood stands under different silvicultural regimes

Abstract
Pathways of forest compositional dynamics over five decades (1953-2005) were reconstructed using measurements from permanent sample plots in a long-term silviculture experiment in a central Maine mixedwood forest. The objective of this study was to elucidate the dynamics of tree species composition at the sample plot level in relation to the initial composition when the experiment was established (1953-1957) and harvest disturbance history following a wide range of even-aged and uneven-aged silvicultural treatments. Cluster analysis revealed three groupings of sample plots based on pretreatment composition and harvest disturbance history, or nine subclusters (i.e., three harvest disturbance histories nested within each pretreatment composition). From 1953 to 2005, the silvicultural treatments generated an array of compositional outcomes at the plot level. Hardwood dominance increased following a history of heavy and infrequent harvests, while northern conifer dominance was maintained where harvests were lighter and more frequent. The importance of balsam fir (Abies balsamea (L.) Mill.) changed little across a range of harvest intensities. A ubiquitous decline in northern white-cedar (Thuja occidentalis L.) was found among silvicultural treatments, suggesting that additional intervention may be needed to promote cedar recruitment. Plot-level compositional dynamics indicated that neighborhood-scale stand dynamics were associated with variability in harvest disturbance overlain on plot-to-plot variability in tree species composition at the time the experiment was established.