Haustorial Synergids in Danthonioid Grasses

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Abstract
Synergids with a haustorial projection, which extends through the micropyle to lie between the ovule and ovary wall, are described for 7 danthonioid genera: Chionochloa [rigida], Danthonia [sericea, spicata], Erythranthera [pumila], Lamprothyrsus [peruvianus], Pyrrhanthera [exigua], Rytidosperma [setifolium] and Sieglingia [decumbens]; they were already known in Cortaderia. Normal pyriform graminoid synergids were found in the arundinoid genera Arundo [donae] and Phragmites [australis]. The distribution of this character among genera is in accord with the traditional tribal arrangements of the grasses, except for Cortaderia, which now seems better treated as danthonioid rather than arundinoid. Synergids that extend as haustoria are interpreted as derived from the normal graminoid type even though they occur, so far, in old, highly polyploid genera.

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