IV. Further experimental note on the correlation of action of antagonistic muscles

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Abstract
Appropriate excitation of the afferent nerves from the flexor muscles of the knee joint so alters, as I have shown, the condition of the extensor muscles of that joint that the reaction called the “knee jerk” becomes no longer elicitable. I have endeavoured to examine the quality of the alteration which thus restrains or abolishes the “jerk.” It must be remembered that there is some variance of opinion as to the nature of the jerk itself. In the opinion of some authorities the jerk is of reflex nature (Bowditch, Lombard, Senator, Warren); in the opinion of others it is not truly reflex, but is a direct muscular reaction, intimately dependent, however, on a reflex tonus in the muscle (Tschiriew), or on a spinal influence reflexly exerted, but not necessarily identical with “tonus” nor necessarily measurable by tonicity (Waller).

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