The oldest bovid from the Siwaliks, Pakistan

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Abstract
Two new small-sized bovid species are described from the Miocene of Pakistan (Kamlial and Vihowa formations, about 18 Ma), based on associated dentitions and horn cores. Eotragus noyei, sp. nov. is probably ancestral to all younger Eotragus and the modern Boselaphini. E. noyei has a horn core which does not have a pronounced anterior concavity and posterior convexity seen in younger Eotragus species and it has primitive teeth with inclined buccal walls. The indeterminate genus and species from the Vihowa Formation has compressed, keeled horn cores and more advanced teeth with vertical buccal walls. The morphology of the horn cores suggests that this taxon could represent the earliest known member of the keeled Boselaphini. If this hypothesis is correct, we are witnessing the presence in Pakistan of the first split within the Boselaphini (keeled Miocene species versus Eotragus and modern taxa).