An Ophiceratid ammonoid from the New Zealand Permian and its implications for the Permian–Triassic boundary

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Abstract
A nesw Ophiceratid genus and species Durvilleoceras woodmani is described from the late Middle Permian Greville Formation of New Zealand, with Episageceras aff noetlingi Haniel. The horizon is close in age to the Kathwai dolomite of the Salt Range, which also has a mid-Permian Ophiceratid species, Ophiceras connectens Schindewolf. The occurrence of these two species shows that the Ophiceratidae cannot serve as an index of early Triassic rocks. The so-called basal Triassic Griesbachian Stage may prove to be closely linked to the Permian period. The key Griesbachian ammonoid genus Otoceras is related to Permian genera, with no later survivors. Moreover the widespread occurrence of Permian-type Productacea and other brachiopods in beds of the Griesbachian Stage in North America and Himalayas also suggests that the Griesbachian is Permian rather than Triassic. To judge from faunas, the start of the Triassic could be based on the incoming of numerous ammonoid families and Triassic brachiopods in the Smithian Stage. The intervening Dienerian Stage between the Griesbachian and Smithian stages is relatively barren of faunas, reflecting some sort of catastrophe at the end of the Paleozoic Era, but has mainly Permian survivors. Such a picture of the Paleozoic–Mesozoic boundary conforms with the intention if not the practice of early paleontologists.