Illusory contours: Toward a neurally based perceptual theory

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Abstract
Although illusory contours were first described nearly a century ago, researchers have only recently begun to approach a consensus on the processes underlying their formation. Neurophysiological and psychophysical evidence indicate that neural mechanisms of the early visual cortex subserve illusory contour generation, although cognitive factors play important roles in determining the final percept. I summarize experiments concerning the determinants of illusory contour strength and form, concentrating on findings particularly relevant to modeling. After establishing arguments for the early generation of illusory contours, I provide an overview of formation theories, culminating with descriptions of neural models. The constraints that experimental data place on models are outlined, and neural models are evaluated with respect to these constraints. Throughout the review, I indicate where further experimental and modeling research are critical.