Measurements of electron temperature made in Ariel I have been analyzed to calculate the ionospheric energy input required to maintain the electron temperature above the ion temperature. The results are found to be consistent with the energy input due to photo-ionization in the daytime, provided that allowance is made for the effects of the escaping flux of photoelectrons spiralling upwards along the geomagnetic field lines which impartenergy to the ionosphere by electron-electron interaction. However, it is found that during the night an energy input of particle origin is observed, a close agreement being found between the distribution of energy input and that of the fluxes of low-energy particles observed by Savenko, Shavrin & Pisavenko 1963. The particle flux contributes less than 30% to the heat input in the daytime and its diurnal variation is small.