Spatial Heterogeneity and Bird Species Diversity

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Abstract
A heterogeneity index, D, derived from the point—quarter technique was significantly correlated with bird species diversity (BSD) for several shrub and forest areas. It predicted BSD for a series of similar brushlands where other indices had failed. Species richness increased faster than species overlap in a series of increasing complex habitats up to the forest stage. Species overlap was negatively correlated with patchiness. Additional species may be accommodated in preforest habitats primarily by horizontal spatial segregation facilitate by the presence of additional patches. New patches result from the addition of layers of vegetation. In late shrub or forest stages other kinds of segregation such as vertical segregation become important to species packing. Patchiness, as measured here, has a proximate effect on avian diversity. The extent or existence of latitudinal gradients in habitat patchiness and the effect on bird species diversity is unknown. The need remains for a universal simple, yet meaningful, heterogeneity index which incorporates both horizontal and vertical variability of vegetation.