Proline transport and oxidation in short-circuited locust rectum: effect of cAMP

Abstract
Proline absorption from the lumen is the main source of substrate supporting electrogenic Cl transport and short-circuit current (Isc) in locust rectum. Since cAMP stimulates chloride-dependent Isc by up to 10-fold, we investigated whether this stimulant also increases active transport of the major metabolic substrate proline. A large 40:1 flux ratio of [14C]proline under short-circuited conditions confirmed that net absorption of this amino acid is by active transport. Unexpectedly, cAMP caused a 40% decrease in net transepithelial flux of proline. A 45% increase in the amount of proline oxidized to 14CO2 could account for only 10% of this decrease in proline flux, suggesting that increased Cl transport after stimulation may competitively reduce the energy supply for proline transport.

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