A molecular approach to the study of genic heterozygosity in natural populations. 3. Direct evidence of coadaptation in gene arrangements of Drosophila.

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Abstract
Classification of electro-phoretically separable alleles at 2 loci on the 3rd chromosome of D. pseudoobscura and D. persimilis was correlated with inversion poly-morphism on the chromosome. Different inversions are genetically differentiated and the differentiation has been maintained over several million generations by natural selection.