Seed Morphology in the Tribe Morindeae (Rubiaceae)

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Abstract
The New World genus Appunia and New World representatives of Morinda of the tribe Morindeae (Rubiaceae) have unique fruits and seeds which readily segregate these from 41 genera of the Rubioideae. The fruits are 4-loculate, each locule with one pyrene. The endocarp of the pyrene develops a bladder-like cavity on the adaxial side. The seeds are vascular and winged; the trabeculae of the alar cells readily segregate the genera. The sclerenchyma elements of the seed suggest a close relationship between the tribe Morindeae and the tribe Coussareae. The description of the Panamanian species, Appunia seibertii Standley, used in the present study, is emended; a key to the Central American spp. of Morinda and Appunia is included.

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