Revision of the Genus Cochlidium (Grammitidaceae)

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Abstract
The genus includes neotropical, grammitid ferns with simple and entire, or at most, sinuate fertile laminae, concolorous scales, hydathodes, and 2-8-celled hairs which are characterized by thickened intercellular walls and are frequently catenate. Two of the species, C. rostratum and C. linearifolium are highly derived and glabrous. Another unifying and distinguishing feature is the stelar organization which shows 1-3 vascular bundles, each with a straight or lightly arched row of tracheids instead of the usual family arrangement in a solenostele. Also, some of the species have a coenosorus, and a tendency of the fertile lamina to fold conduplicately over the sporangia. New taxa or combinations proposed are: C. serrulatum comb. nov. (Acrostichum serrulatum Swartz); C. punctatum comb. nov. (Grammitis punctatum Raddi); C. proctorii comb. nov. (Grammitis proctorii Copel.); C. tepuiense comb. nov. (Polypodium tepuiense A.C. Smith); C. repandum sp. nov. (Arroyo del Medio, Sierra de Nipe, Oriente, Cuba, 450-500 m on tree fern bases above the falls and in eastern Cuba north to Baracoa); C. jungens sp. nov. (epiphytic in mossy forest, top of Morne Micotrin, St. George Parish, Dominica, and Puerto Rico, 800-1500 m); and C. wurdackii sp. nov. (river banks, mats on rocks, Sarven-tepui, Edo. Bolivar, Venezuela, 1200 m). Pleurogramme myrtillifolia is excluded pending rediscovery of the type. [A key to the following species was also provided: C. seminudum, C. graminoides, C. pumilum, C. minus, C. furcata and C. dubius.].

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