IV. On the relations of the nucleus to spore-formation in certain liver-worts

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Abstract
It is well known that, as a general rule, during the formation of spore tetrads from their mother cells, the nucleus of the latter commonly undergoes two successive bipartitions. Each of the resulting four nuclei ultimately becomes a centre for the aggregation of a portion of the original protoplasm, whilst division of the whole immediately follows by means of cell walls. Though the above method is the one most commonly followed during the process of spore development, so far as the essential features are concerned, it is by no means the invariable one.

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