Abstract
We present an analysis of the cosmological evolution of matter sources with small anisotropic pressures. This includes electric and magnetic fields, collisionless relativistic particles, gravitons, antisymmetric axion fields in low-energy string cosmologies, spatial curvature anisotropies, and stresses arising from simple topological defects. We calculate their evolution during the radiation and dust eras of an almost isotropic universe. In many interesting cases the evolution displays a special critical behaviour created by the non-linear evolution of the pressure and expansion anisotropies. The isotropy of the microwave background is used to place strong limits of order $\Omega _{a0}\leq 5\times 10^{-6}\Delta (1+z_{rec})^{-\Delta }$on the possible contribution of these matter sources to the total density of the universe, where $1\leq \Delta \leq 3$ characterises the anisotropic stress. The present abundance of an anisotropic stress which becomes non-relativistic at a characteristic low-energy scale is also calculated. We explain why the limits obtained from primordial nucleosynthesis are generally weaker than those imposed by the microwave background isotropy. The effect of inflation on these stresses is also calculated.

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