A field study of settlement behaviour inBalanus balanoidesandElminius modestus(Cirripedia: Crustacea) in relation to competition between them

Abstract
There can be little doubt that competition for space, both inter- and intraspecific is a major factor in barnacle ecology. Southward & Crisp (1956) first pointed out the role of competition betweenBalanus balanoides(L.) andChthamalus stellatus(Poli) in south-west Britain which Connell (1959, 1961) later demonstrated experimentally. Crisp (1958) discussed competition between the indigenousB. balanoidesand the immigrantElminius modestusDarwin in the intertidal zone, and measurements of growth rates, using manipulated populations of these two species, indicated that inter- and intraspecific competition for space after settlement is intense (Crisp, 1964).
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