SYSTEMATICS OF SHUMARDIIDAE (TRILOBITA), WITH NEW SPECIES FROM THE ORDOVICIAN OF ARGENTINA

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Abstract
New shumardiid trilobites from Arenig to Early Caradoc deposits of the Argentine Precordillera include species of a revised Kweichowilla (K. salasae, K. sterrenae, and K. piojensis) and the new genus Changchowilla (C. sanjuanina and C. gracielae, and C.? carrerai). Conophrys fabiani new species occurs in the Early Tremadoc of the western Puna region, northwestern Argentina. Leoforteyia new genus is based on L. hintzei new species from the Ibexian of Utah. Published taxa recognized as new species are Conophrys rushtoni, Conophrys wrighti, Leoforteyia ludvigseni and Shumardia whittingtoni. Phylogenetic analysis suggests the differentiation of Shumardia Billings, 1862; Kweichowilla Chang, 1964; Changchowilla new genus; and Leoforteyia new genus as separate taxa and indicates that Conophrys Callaway, 1877, is a grade group. Silicified material provides a nearly complete growth series for Kweichowilla salasae, including the first protaspides known with confidence for the Shumardiidae. Additionally, on...